Ho Ting-nong、Liu Shang-wu编著的《獐牙菜属和近缘属的世界性分类修订英文版》介绍了,獐牙菜属及近缘植物中獐牙菜苦苷、龙胆苦苷、芒果苷、齐墩果酸的含量,为资源利用提供依据。獐牙菜属(Swertia)是龙胆科下的一个属,约100种,广布于全球,我国约70余种,南北均产之。一年生或多年生草本。一年生或多年生草本;叶对生;花蓝色或白色,有黄色的腺点,排成密锥花序式或伞房花序式的圆锥花序;萼片4-5深裂,每一裂片的近基部有窝孔或腺体1-2个,窝孔秃裸或一部为一基生的鳞片所覆盖,其周边或顶端具睫毛;雄蕊4-5,着生于花冠的近基部,分离,很少合生成1束;子房1室,柱头2;果为一卵形或长椭圆形的蒴果,成熟时分裂为2果瓣,有种子多数。
目錄:
Contents
1 Inter-generic circumscription of Swertia and allies1
1.1An introduction to the tribe Gentianae1
1.2Key to genera of the tribe Gentianae and subtribe Swertiinae1
1.3Morphologica ldifferentiation between the rotate and tubular genera4
2 SwertiaLinnaeus12
2.1Historical survey12
2.2Generic circumscription in the present work18
2.2.1Frasera21
2.2.2Ophelia22
2.2.3Anagallidium22
2.3Taxonomic traits of Swertia sensu lato23
2.3.1Grossmorphology23
2.3.2Embryology38
2.3.3Chromosomes44
2.3.4Pollentraits47
2.3.5Pollen traits of subgenera and sections68
2.3.6Taxonomic implications of pollen traits83
2.4Distribution and diversification85
2.5Taxonomic treatment87
2.5.1Anenumeration of SwertiaL.88
2.5.2Keytotaxa94
2.5.3Asystematic account114
2.6Unknown uncertain species299
2.7The names excluded from SwertiaL.300
3 SinoswertiaT.N.Ho,S.W.Liu&J.Q.Liu303
3.1Taxonomic treatments303
3.2Reproductive biology306
3.3Embryology307
3.4Systematic position308
4 LomatogoniopsisT.N.Ho&S.W.Liu309
4.1Taxonomic treatment309
4.2Pollen morphology313
4.3Systematic position314
5 Veratrilla Baillonex Franchet315
5.1Taxonomic treatment315
5.2Pollen morphology318
5.3Systematic position318
6 Lomatogonium A.Braun319
6.1Taxonomic treatment319
6.2Embryology350
6.3Chromosome numbers351
6.4Pollen morphology352
6.5Uncertain species354
6.6The names excluded from Lomatogonium355
7 Mainreferences361
8 Representative specimens373
8.1Representative specimens for Swertia Linnaeus373
8.2Representative specimens for SinoswertiaT.N.Ho,S.W.Liu&J.Q.Liu420
8.3Representative specimens for LomatogoniopsisT.N.Ho&S.W.Liu fu hua421
8.4Representative specimens for VeratrillaBaillon ex Franchet422
8.5Representative specimensfor LomatogoniumA.Braun423
8.6Index to collectors of cited specimens431
Index465
內容試閱:
1 Inter-generic circumscription of Swertia and allies
1.1 An introduction to the tribe Gentiana
The tribe Gentiana circumscribed by Struwes and Albert 2002 comprises 17 genera in two subtribes. Subtribe Gentianinae contains Gentiana, Crawfurdia, Tripterospermum, while subtribe Swertiinae includes Bartonia, Comastoma, Frasera, Gentianella, Gentianopsis Halenia, Jaeschkea, Latouchea, Lomatogonium, Megacodon, Obolaria, Pterygocalyx, Swertia and Veratrilla. Within the past three decades, we published three additional genera to be included in the Gentianae Lomatogoniopsis, Metagentiana and Sinoswertia Ho and Liu, 1980; Ho et al., 2002; He et al., 2013b. Phylogenetic analyses constructed based on molecular evidence suggested that after erecting Sinoswertia, the genus Swertia sensu lato was still polyphyletic Yuan and Küpfer 1995; Chassot et al., 2001; Liu et al., 2001; He et al., 2013b. However, a limited number of Swertia species were sampled. It remains elusive how many genera should be recognized within Swertia sensu lato. Here, we tentatively adopt the generic circumscriptions of most genera described by previous authors for example, Struwes and Albert, 2002. In addition, we further place Frasera in Swertia sensu lato based on available morphological comparisons. Thus, the Gentianae should comprise 19 genera in total according to current taxonomic treatments. Both subtribes Gentianinae and Swertiinae can be easily distinguished by the presence or lack of corolla plicae and intracalycular membranes between lobes Smith, 1936. The subtribe Swertiinae also may be subdivided into two groups: the Rotate group and the Tubular group Smith, 1965. The former has a rotate corolla and calyx, with nectaries either naked or surrounded by fringed or tubular appendages, whereas the latter has a tubular corolla and calyx, with nectaries naked or absent Ho and Liu, 2001. In the present book, we present a monograph of the rotate group comprising Swertia, Sinoswertia, Lomatogoniopsis, Lomatogonium, and Veratrilla.
1.2 Key to genera of the tribe Gentiana and subtribe Swertiinae
1a. Corolla with plicae extending between lobes except in G. lutea L.; with an intracalycular membrane between lobes except in Crawfurdia spp. and Tripterospermum spp. subtribe Gentianinae:
2a. Flowers ebracteate at base except for a few species in sections Pneumonanthe and Gentiana; style linear to cylindrical, shorter than ovary; seeds ellipsoid, ovoid to global, not triquetrous; calyx tube with 15 vascular bundlesGentiana L.
2b. Flowers bracteate at base; style filiform, as long or longer than ovary; seeds triquetrous to compressed, with three edges; calyx tube with 5–12 vascular bundles:
3a. Stems striate-angled, erect, neither twining nor trailing; flowers sessile, solitary and terminal at branches; bracts of flowers as large or larger than stem leaves; fruit included in persistent corolla; seeds triquetrous with narrow wings on edges, three edges forming three equal faces; annuals, rarely perennials with stout and extremely shortened disc-like rhizome
Metagentiana T. N. Ho and S. W. Liu
3b. Stems terete, twining or trailing; flowers 1–5, pedicellate, in terminal and axillary cymes; bracts of flowers much smaller than stem leaves; fruit exserted from persistent corolla; seeds compressed with discoid wings on edges, rarely triquetrous to compressed with narrow wings, three seed edges forming three extremely unequal faces; perennials with stout, long and branched rhizomes:
4a. Nectaries conspicuously developed, forming a collarlike disc around gynophore; stamens, unequal, apically decurved; fruit a capsule or berry; calyx tube with 5 vascular bundlesTripterospermum Blume
4b. Nectaries small, on gynophore; stamens symmetrical, equal, straight; fruits a capsule; calyx tube with 10 vascular bundlesCrawfurdia Wallich
1b. Corolla without plicae between lobes; without intracalycular membrane between lobes except Gentianopsis subtribe Swertiinae:
5a. Corolla rotate, lobed nearly to base, tube very short, indistinct, much shorter than lobes; nectaries surrounded by fringed or tubular appendages or naked:
6a. Plants dioecious, with unisexual flowers; seeds discoid-winged
Veratrilla Baillon ex Franchet
6b. Plants with perfect flowers; seeds various:
7a. Nectaries not surrounded by fringes or appendagesGentianella Moench
7b. Nectaries surrounded by fringed or tubular appendages:
8a. Both plants and flowers dimorphic: plants heterogeneous and homog-eneous; the former with a distinct main stem and numerous weak branches at stem base, on main stem flowers larger and nectaries linear-oblong fringed, on weak branches flowers 14–13 as large as those of main stem and nectaries with broadly oblong entire elevated scales and indistinct flat gland patches; the latter with uniform stems and flowers, flowers are the same in shape, size and nectaries as those of large-sized flowers on heterogeneous plants
Sinoswertia T. N. Ho, S. W. Liu and J.